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doi: 10.1038/nature11260

Intracellular antioxidant enzymes function to reduce ROS levels

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This can happen for a variety of reasons, including tobacco that has been dropped on the floor, broken up into small pieces, discolored, dirty, stems, etc

Zafeiridou, M., Hopkinson, N

Forde M., Forde SCF., Disaster Preparedness and Response Plan For HIV/AIDS Interventions In Emergencies In Grenada, The United Nations Trust Fund for Human Security (UNTFHS) Project, May 2009