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286,287 Among the SCFAs, butyrate is the most potent HDAC inhibitor that is generally thought to inhibit the activity of class I HDACs (HDAC1, 2, 3, and 8) and class IIa HDACs (HDAC4, 5, 7, and 9), but not class IIb HDACs (HDAC6 and HDAC10) and class III HDACs (sirtuins)