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Acknowledgements The authors thank Nick Rhodes from the Queensland Cyber Infrastructure Foundation (QCIF), Haojing Shao, and Enya Hermsen for their advice on bioinformatical analyses and computational assistance, Alastair McEwan and Mathias Jnsson for their editorial assistance on early drafts of the manuscript, and all members of the Degnan lab for their valuable discussions

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67 Non-proliferative astrogliosis typically occurs in neural tissue that is responding to a pathology but is not overtly damaged and retains its basic tissue architecture, for example (i) during diffuse neuroinflammation caused by peripheral exposure to microbial antigens such as lipopolysaccharide (LPS), 68,69 (ii) in tissue regions that are at a distance from focal lesions (diaschisis) caused by stroke, trauma or autoimmune attack, 8,70 or (iii) in tissue undergoing neurodegenerative changes

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