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Further, cytochrome c is released as the outer mitochondrial membrane is destroyed, due to loss of mitochondrial membrane potential, which can be diminished by treatment with mitochondrial respiration inhibitors, such as oligomycin, to initiate apoptosis even before apoptotic morphological features appear (69)

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The treatment effectively restored antioxidant defenses by increasing GSH and CAT activity while significantly reducing MDA levels

doi: 10.1016/S0223-5234(03)00082-5 60 Von GruenigenVEJamisonJMGilloteauxJLorimerHESummersMPollardRRet al