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The biosynthesis of diterpenes involves the formation of the precursor geranylgeranyl diphosphate (GGPP), catalyzed by GGPP synthase, followed by the formation of a carbon backbone, which is catalyzed by diterpene synthases (DTSs), and further structural modifications catalyzed by cytochrome P 450 oxygenases (CPYs or P450s) (Table 2)

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doi:10.1016/j.thromres.2009.02.010 Sobenin IA, Myasoedova VA, Iltchuk MI, Zhang DW, Orekhov AN